1. Microbiology Department, School of Natural Sciences, National University of Ireland, Galway, Ireland
  2. Information and Computational Sciences, James Hutton Institute, Invergowrie, Dundee DD2 5DA, Scotland
  3. Soil and Environmental Microbiology, Environmental Research Centre, Teagasc, Johnstown Castle, Wexford, Ireland
  4. School of Life Sciences, University of Bedfordshire, Luton, UK