Comparing the metabolism of C. hyperboreus at depth and at the surface in early fall near Spitsbergen, Auel et al. (2003) found a reduced metabolism of the deep individuals (probably already in diapause) compared the surface ones. The same observation has been done by Darnis et al. 2012 in the Amundsen Gulf, where all the metazoans bigger than 1000µm have a respiration 4 to 5-fold different between the spring values and the late winter ones. From there, we will assume that the two main parameters influencing the accumulation and the use of these lipids are respectively the nitrogen excretion and the respiration.