But the scope of the analogy between evolution and non-equilibrium physics is, in fact, much broader: the interplay between selection and mutation is typical of localization phenomena in disordered systems \cite{Stollmann_2001}, be them classical or quantum. The linearized Crow-Kimura equation \ref{PAM}, for instance, is formally identical to the parabolic Anderson model  \cite{Zel_dovich_1987,Carmona_1994,K_nig_2016}, a simple model of intermittency in random fluid flows; the linearized Eigen model in turn resembles the Bouchaud trap model \cite{Bouchaud_1992}, a classical model of slow dynamics and ageing in glassy systems. These physical phenomena have obvious evolutionary counterparts: the Anderson localization transition corresponds to the error threshold; intermittency to epochal or punctuated evolution; tunnelling instantons to fitness valley crossings; and ageing to diminishing-return epistasis. The generalization of Nelson's mapping of the Scrödinger equation to a diffusion process presented in this paper implies that all are in fact unified under the familiar umbrella of Markovian metastability.