Muller, R. U. & Kubie, J. L. The firing of hippocampal place cells predicts the future position of freely moving rats. J. Neurosci.  9 , 4101–4110 (1989)
Nauta, W. J. H., & Kuypers, H. G. J. M. (1958). Some ascending pathways in the brain stem reticular formation. In H. H. Jasper, L. D. Proctor, R. S. Knighton, W. C. Noshay, & R. T. Costello (Eds.), Reticular formation of the brain  (pp. 3-30). Oxford, England: Little, Brown.
Newman, E. L., Gillet, S. N., Climer, J. R., & Hasselmo, M. E. (2013). Cholinergic blockade reduces theta-gamma phase amplitude coupling and speed modulation of theta frequency consistent with behavioral effects on encoding. Journal of Neuroscience33 (50), 19635-19646.
Niell, C. M., & Stryker, M. P. (2010). Modulation of visual responses by behavioral state in mouse visual cortex. Neuron65 (4), 472-479.
Niell, C. M. (2015). Cell types, circuits, and receptive fields in the mouse visual cortex. Annual review of neuroscience38 , 413-431. Nitz, D. A., & McNaughton, B. L. (1999). Hippocampal EEG and unit activity responses to modulation of serotonergic median raphe neurons in the freely behaving rat. Learning & Memory6 (2), 153-167.
Nitz, D., & McNaughton, B. (2004). Differential modulation of CA1 and dentate gyrus interneurons during exploration of novel environments. Journal of neurophysiology91 (2), 863-872.
Nitz, D. A. (2006). Tracking route progression in the posterior parietal cortex. Neuron49 (5), 747-756.
Noga, B. R., Sanchez, F. J., Villamil, L. M., O’Toole, C., Kasicki, S., Olszewski, M., … & Jordan, L. M. (2017). LFP oscillations in the mesencephalic locomotor region during voluntary locomotion. Frontiers in neural circuits11 .
Norton, A. B., Jo, Y. S., Clark, E. W., Taylor, C. A., & Mizumori, S. J. (2011). Independent neural coding of reward and movement by pedunculopontine tegmental nucleus neurons in freely navigating rats. European Journal of Neuroscience33 (10), 1885-1896.
O’Keefe, J., & Dostrovsky, J. (1971). The hippocampus as a spatial map. Preliminary evidence from unit activity in the freely-moving rat. Brain research34 (1), 171-175.
Parker, S. M., & Sinnamon, H. M. (1983). Forward locomotion elicited by electrical stimulation in the diencephalon and mesencephalon of the awake rat. Physiology & behavior .
Parron, C., & Save, E. (2004). Evidence for entorhinal and parietal cortices involvement in path integration in the rat. Experimental Brain Research159 (3), 349-359.
Patel, J., Fujisawa, S., Berényi, A., Royer, S., & Buzsáki, G. (2012). Traveling theta waves along the entire septotemporal axis of the hippocampus. Neuron75 (3), 410-417.
Patterson, C. M., Wong, J. M. T., Leinninger, G. M., Allison, M. B., Mabrouk, O. S., Kasper, C. L., … & Myers Jr, M. G. (2015). Ventral tegmental area neurotensin signaling links the lateral hypothalamus to locomotor activity and striatal dopamine efflux in male mice. Endocrinology156 (5), 1692-1700.
Perez-Escobar, J. A., Kornienko, O., Latuske, P., Kohler, L., & Allen, K. (2016). Visual landmarks sharpen grid cell metric and confer context specificity to neurons of the medial entorhinal cortex. Elife5 , e16937.
Peyrache, A., Lacroix, M. M., Petersen, P. C., & Buzsáki, G. (2015). Internally organized mechanisms of the head direction sense. Nature neuroscience18 (4), 569.
Pinto, L., Goard, M. J., Estandian, D., Xu, M., Kwan, A. C., Lee, S. H., … & Dan, Y. (2013). Fast modulation of visual perception by basal forebrain cholinergic neurons. Nature neuroscience16 (12), 1857-1863.
Puryear, C. B., Kim, M. J., & Mizumori, S. J. (2010). Conjunctive encoding of movement and reward by ventral tegmental area neurons in the freely navigating rodent. Behavioral neuroscience124 (2), 234.
Quirk, G. J., Muller, R. U., Kubie, J. L., & Ranck, J. B. (1992). The positional firing properties of medial entorhinal neurons: description and comparison with hippocampal place cells. Journal of Neuroscience12 (5), 1945-1963. Ray, S., & Maunsell, J. H. (2010). Differences in gamma frequencies across visual cortex restrict their possible use in computation. Neuron67 (5), 885-896.
Reifenstein, E. T., Ebbesen, C. L., Tang, Q., Brecht, M., Schreiber, S., & Kempter, R. (2016). Cell-type specific phase precession in layer II of the medial entorhinal cortex. Journal of Neuroscience36 (7), 2283-2288.
Richard, G. R., Titiz, A., Tyler, A., Holmes, G. L., Scott, R. C., & Lenck‐Santini, P. P. (2013). Speed modulation of hippocampal theta frequency correlates with spatial memory performance. Hippocampus23 (12), 1269-1279.
Rivas, J., Gaztelu, J. M., & Garcia-Austt, E. (1996). Changes in hippocampal cell discharge patterns and theta rhythm spectral properties as a function of walking velocity in the guinea pig. Experimental Brain Research108 (1), 113-118.
Robinson, J., Manseau, F., Ducharme, G., Amilhon, B., Vigneault, E., El Mestikawy, S., & Williams, S. (2016). Optogenetic activation of septal glutamatergic neurons drive hippocampal theta rhythms. Journal of Neuroscience36 (10), 3016-3023.
Ropert, N. (1985). Modulation of inhibition in the hippocampus in vivo. Canadian journal of physiology and pharmacology63 (7), 838-842.
Roseberry, T. K., Lee, A. M., Lalive, A. L., Wilbrecht, L., Bonci, A., & Kreitzer, A. C. (2016). Cell-type-specific control of brainstem locomotor circuits by basal ganglia. Cell164 (3), 526-537.
Roth, M. M., Dahmen, J. C., Muir, D. R., Imhof, F., Martini, F. J., & Hofer, S. B. (2016). Thalamic nuclei convey diverse contextual information to layer 1 of visual cortex. Nature neuroscience19 (2), 299-307.
Ryczko, D., & Dubuc, R. (2013). The multifunctional mesencephalic locomotor region. Current pharmaceutical design19 (24), 4448-4470.
Saleem, A. B., Ayaz, A., Jeffery, K. J., Harris, K. D., & Carandini, M. (2013). Integration of visual motion and locomotion in mouse visual cortex. Nature neuroscience16 (12), 1864-1869.
Sanes, J. N., & Donoghue, J. P. (1993). Oscillations in local field potentials of the primate motor cortex during voluntary movement. Proceedings of the National Academy of Sciences90 (10), 4470-4474.
Sargolini, F., Fyhn, M., Hafting, T., McNaughton, B. L., Witter, M. P., Moser, M. B., & Moser, E. I. (2006). Conjunctive representation of position, direction, and velocity in entorhinal cortex. Science312 (5774), 758-762.
Scaplen, K. M., Ramesh, R. N., Nadvar, N., Ahmed, O. J., & Burwell, R. D. (2017). Inactivation of the lateral entorhinal area increases the influence of visual cues on hippocampal place cell activity. Frontiers in systems neuroscience11 .
Schneider, D. M., Nelson, A., & Mooney, R. (2014). A synaptic and circuit basis for corollary discharge in the auditory cortex. Nature513 (7517), 189-194.
Segal, M. (1978). A correlation between hippocampal responses to interhemispheric stimulation, hippocampal slow rhythmic activity and behaviour. Electroencephalography and clinical neurophysiology45 (3), 409-411.
Sharp, P. E., Blair, H. T., & Cho, J. (2001). The anatomical and computational basis of the rat head-direction cell signal. Trends in neurosciences24 (5), 289-294.
Shen, J., Barnes, C. A., McNaughton, B. L., Skaggs, W. E., & Weaver, K. L. (1997). The effect of aging on experience-dependent plasticity of hippocampal place cells. Journal of Neuroscience17 (17), 6769-6782.
Sheremet, A., Burke, S. N., & Maurer, A. P. (2016). Movement enhances the nonlinearity of hippocampal theta. Journal of Neuroscience36 (15), 4218-4230.
Shik, M. L., Severin, F. V., & Orlovskiĭ, G. N. (1966). Control of walking and running by means of electric stimulation of the midbrain. Biofizika11 (4), 659-666.
Shin, J., & Talnov, A. (2001). A single trial analysis of hippocampal theta frequency during nonsteady wheel running in rats. Brain research897 (1), 217-221.
Singer, W. (1999). Neuronal synchrony: a versatile code for the definition of relations?. Neuron24 (1), 49-65. Sirota, A., Montgomery, S., Fujisawa, S., Isomura, Y., Zugaro, M., & Buzsáki, G. (2008). Entrainment of neocortical neurons and gamma oscillations by the hippocampal theta rhythm. Neuron60 (4), 683-697.
Skinner, R. D., & Garcia-Rill, E. (1984). The mesencephalic locomotor region (MLR) in the rat. Brain research323 (2), 385-389.
Sławińska, U., & Kasicki, S. (1998). The frequency of rat’s hippocampal theta rhythm is related to the speed of locomotion. Brain research796 (1), 327-331.
Smythe, J. W., Colom, L. V., & Bland, B. H. (1992). The extrinsic modulation of hippocampal theta depends on the coactivation of cholinergic and GABA-ergic medial septal inputs. Neuroscience & Biobehavioral Reviews16 (3), 289-308.
Strange, B. A., Witter, M. P., Lein, E. S., & Moser, E. I. (2014). Functional organization of the hippocampal longitudinal axis. Nature Reviews Neuroscience15 (10), 655-669.
Sotty, F., Danik, M., Manseau, F., Laplante, F., Quirion, R., & Williams, S. (2003). Distinct electrophysiological properties of glutamatergic, cholinergic and GABAergic rat septohippocampal neurons: novel implications for hippocampal rhythmicity. The Journal of physiology551 (3), 927-943.
Stackman, R. W., Clark, A. S., & Taube, J. S. (2002). Hippocampal spatial representations require vestibular input. Hippocampus12 (3), 291-303.
Stewart, D. J., & Vanderwolf, C. H. (1987). Hippocampal rhythmical slow activity following ibotenic acid lesions of the septal region. I. Relations to behavior and effects of atropine and urethane. Brain research423 (1), 88-100.
Sun, Y., Nguyen, A. Q., Nguyen, J. P., Le, L., Saur, D., Choi, J., … & Xu, X. (2014). Cell-type-specific circuit connectivity of hippocampal CA1 revealed through Cre-dependent rabies tracing. Cell reports7 (1), 269-280.
Sun, C., Kitamura, T., Yamamoto, J., Martin, J., Pignatelli, M., Kitch, L. J., … & Tonegawa, S. (2015). Distinct speed dependence of entorhinal island and ocean cells, including respective grid cells. Proceedings of the National Academy of Sciences112 (30), 9466-9471.
Swanson, L. W., & Cowan, W. M. (1979). The connections of the septal region in the rat. Journal of Comparative Neurology186 (4), 621-655.
Takakusaki, K. (2008). Forebrain control of locomotor behaviors. Brain research reviews57 (1), 192-198.
Tanke, N., Carvalho, M. M., Kropff Causa, E., Witter, M. P., Moser, M-B, & Moser, E. I. (Unpublished results). A brainstem/basal forebrain/cortical circuit for the neuronal coding of locomotion speed. In Society for Neuroscience Abstracts, Program (Neuroscience 2017).
Taube, J. S., Muller, R. U., & Ranck, J. B. (1990a). Head-direction cells recorded from the postsubiculum in freely moving rats. I. Description and quantitative analysis. Journal of Neuroscience10 (2), 420-435.
Taube, J. S., Muller, R. U., & Ranck, J. B. (1990b). Head-direction cells recorded from the postsubiculum in freely moving rats. II. Effects of environmental manipulations. Journal of Neuroscience10 (2), 436-447.
Taube, J. S. (2007). The head direction signal: origins and sensory-motor integration. Annu. Rev. Neurosci.30 , 181-207.
Terrazas, A., Krause, M., Lipa, P., Gothard, K. M., Barnes, C. A., & McNaughton, B. L. (2005). Self-motion and the hippocampal spatial metric. Journal of Neuroscience25 (35), 8085-8096.
Tóth, K., Freund, T. F., & Miles, R. (1997). Disinhibition of rat hippocampal pyramidal cells by GABAergic afferents from the septum. The Journal of physiology500 (2), 463-474.
Traub, R. D., Jefferys, J. G., & Whittington, M. A. (1999). Fast oscillations in cortical circuits . MIT press.
Tsao, A., Sugar, J., Lu, L., Wang, C., Knierim, J. J., Moser, M. B., & Moser, E. I. (2018). Integrating time from experience in the lateral entorhinal cortex. Nature561 (7721), 57.
Unal, G., Joshi, A., Viney, T. J., Kis, V., & Somogyi, P. (2015). Synaptic targets of medial septal projections in the hippocampus and extrahippocampal cortices of the mouse. Journal of Neuroscience35 (48), 15812-15826.
Vandecasteele, M., Varga, V., Berényi, A., Papp, E., Barthó, P., Venance, L., ... & Buzsáki, G. (2014). Optogenetic activation of septal cholinergic neurons suppresses sharp wave ripples and enhances theta oscillations in the hippocampus. Proceedings of the National Academy of Sciences111(37), 13535-13540.
Vanderwolf, C. H. (1969). Hippocampal electrical activity and voluntary movement in the rat. Electroencephalography and clinical neurophysiology26 (4), 407-418.
Vinck, M., Batista-Brito, R., Knoblich, U., & Cardin, J. A. (2015). Arousal and locomotion make distinct contributions to cortical activity patterns and visual encoding. Neuron86 (3), 740-754.
von Nicolai, C., Engler, G., Sharott, A., Engel, A. K., Moll, C. K., & Siegel, M. (2014). Corticostriatal coordination through coherent phase-amplitude coupling. Journal of Neuroscience34 (17), 5938-5948.
Wall, N. R., De La Parra, M., Callaway, E. M., & Kreitzer, A. C. (2013). Differential innervation of direct-and indirect-pathway striatal projection neurons. Neuron79 (2), 347-360.
Wang, D. V., & Tsien, J. Z. (2011). Conjunctive processing of locomotor signals by the ventral tegmental area neuronal population. PLoS One6 (1), e16528. Wells, C. E., Amos, D. P., Jeewajee, A., Douchamps, V., Rodgers, J., O’Keefe, J., … & Lever, C. (2013). Novelty and anxiolytic drugs dissociate two components of hippocampal theta in behaving rats. Journal of Neuroscience33 (20), 8650-8667.
Whishaw, I. Q., & Vanderwolf, C. H. (1973). Hippocampal EEG and behavior: Change in amplitude and frequency of RSA (Theta rhythm) associated with spontaneous and learned movement patterns in rats and cats. Behavioral biology8 (4), 461-484.
Whishaw IQ, McKenna JE, Maaswinkel H (1997) Hippocampal lesions and path integration. Curr Opin Neurobiol 7:228 –234. Whishaw, I. Q., Hines, D. J., & Wallace, D. G. (2001). Dead reckoning (path integration) requires the hippocampal formation: evidence from spontaneous exploration and spatial learning tasks in light (allothetic) and dark (idiothetic) tests. Behavioural brain research127 (1), 49-69.
Whishaw, I. Q., & Wallace, D. G. (2003). On the origins of autobiographical memory. Behavioural Brain Research138 (2), 113-119.
Whitlock, J. R., Pfuhl, G., Dagslott, N., Moser, M. B., & Moser, E. I. (2012). Functional split between parietal and entorhinal cortices in the rat. Neuron73 (4), 789-802.
Widmer, H., Ferrigan, L., Davies, C. H., & Cobb, S. R. (2006). Evoked slow muscarinic acetylcholinergic synaptic potentials in rat hippocampal interneurons. Hippocampus16 (7), 617-628.
Wiener, S. I., Paul, C. A., & Eichenbaum, H. (1989). Spatial and behavioral correlates of hippocampal neuronal activity. Journal of Neuroscience9 (8), 2737-2763.
Wilber, A. A., Skelin, I., Wu, W., & McNaughton, B. L. (2017). Laminar organization of encoding and memory reactivation in the parietal cortex. Neuron95 (6), 1406-1419.
Wills, T. J., Barry, C., & Cacucci, F. (2012). The abrupt development of adult-like grid cell firing in the medial entorhinal cortex. Frontiers in neural circuits6 , 21.
Wilson, M. A., & McNaughton, B. L. (1993). Dynamics of the hippocampal ensemble code for space. Science261 (5124), 1055-1059.
Winne, J., Franzon, R., Miranda, A., Malfatti, T., Patriota, J., Mikulovic, S., … & Leão, R. N. (2018). Salicylate induces anxiety‐like behaviour and slow theta oscillation and abolishes the relationship between running speed and fast theta oscillation frequency. Hippocampus .
Winson, J. (1978). Loss of hippocampal theta rhythm results in spatial memory deficit in the rat. Science201 (4351), 160-163.
Winson, J., & Abzug, C. (1978). Neuronal transmission through hippocampal pathways dependent on behavior. Journal of Neurophysiology41 (3), 716-732.
Winter, S. S., Mehlman, M. L., Clark, B. J., & Taube, J. S. (2015). Passive transport disrupts grid signals in the parahippocampal cortex. Current Biology25 (19), 2493-2502. Witter, M.P. and Amaral, D.G. (2004) Hippocampal Formation. In The Rat Nervous System (3rd edn) (Paxinos, G., ed.), pp. 637–703, Elsevier Academic Press
Wolbers, T., Wiener, J. M., Mallot, H. A., & Büchel, C. (2007). Differential recruitment of the hippocampus, medial prefrontal cortex, and the human motion complex during path integration in humans. Journal of Neuroscience27 (35), 9408-9416.
Wyble, B. P., Hyman, J. M., Rossi, C. A., & Hasselmo, M. E. (2004). Analysis of theta power in hippocampal EEG during bar pressing and running behavior in rats during distinct behavioral contexts. Hippocampus14 (5), 662-674.
Yang, F. C., Jacobson, T. K., & Burwell, R. D. (2017). Single neuron activity and theta modulation in the posterior parietal cortex in a visuospatial attention task. Hippocampus27 (3), 263-273.
Ye, J., Witter, M. P., Moser, M. B., & Moser, E. I. (2018). Entorhinal fast-spiking speed cells project to the hippocampus. Proceedings of the National Academy of Sciences115 (7), E1627-E1636.
Yoder, R. M., & Taube, J. S. (2014). The vestibular contribution to the head direction signal and navigation. Frontiers in integrative neuroscience8 , 32.
Yu, J. Y., Kay, K., Liu, D. F., Grossrubatscher, I., Loback, A., Sosa, M., … & Frank, L. M. (2017). Distinct hippocampal-cortical memory representations for experiences associated with movement versus immobility. eLife6 .
Zhang, K., Ginzburg, I., McNaughton, B. L., & Sejnowski, T. J. (1998). Interpreting neuronal population activity by reconstruction: unified framework with application to hippocampal place cells. Journal of neurophysiology79 (2), 1017-1044.
Zheng, C., Bieri, K. W., Trettel, S. G., & Colgin, L. L. (2015). The relationship between gamma frequency and running speed differs for slow and fast gamma rhythms in freely behaving rats. Hippocampus25 (8), 924-938.
Zhou, M., Liang, F., Xiong, X. R., Li, L., Li, H., Xiao, Z., … & Zhang, L. I. (2014). Scaling down of balanced excitation and inhibition by active behavioral states in auditory cortex. Nature neuroscience17 (6), 841-850.
Zhou, T. L., Tamura, R., Kuriwaki, J., & Ono, T. (1999). Comparison of medial and lateral septal neuron activity during performance of spatial tasks in rats. Hippocampus9 (3), 220-234.