The PREreview raised a experimental consideration about sampling with a majority of the isolates/strains coming from California grapes. There are a couple of factors that we feel ameliorates this concern. First, when conducting genomic sequencing, the California grape isolates did not form a distinct population but were instead highly admixed within the entire global population. This suggests that at the genome level the entire collection of strains is behaving as a largely randomized collection. Second, we should have noted that these grapes were all from the UCDavis vineyards that are surrounded by Tomato, Sunflower and Lettuce experimental and production plots. Further, the understory of the vineyard is occuped predominantly by Brassicaceae and Asteraceae weeds.  As such, these isolates have had the chance to be exposed to either the species in our work or at the very least closely related species. We attempted to obtain strains from the understory but the published Botrytis selective media kills a majority of isolates in our hands and we had to proceed with visual identification of the Botrytis.