Supporting Information (optional)

Essential supporting information can be published in the online version of the article. Instructions for the preparation of Supporting Information are given here. Note, however, that the BES does notallow data sets to be uploaded as Supporting Information. All relevant data must be archived in accordance with the BES data archiving policy .
General style points for manuscript preparation
Give Latin names in full at first mention in the main text. Subsequently, the genus name may be abbreviated, except at the beginning of a sentence. If there are many species, cite a Flora or check-list which may be consulted for authorities instead of listing them in the text. Latin names following common names should not be separated by a comma or brackets.
Authors should use the International System of Units (S.I., Système International d’Unités; see Quantities, Units and Symbols, 2nd edn (1975) The Royal Society, London). If the paper contains many symbols, they should be defined as early in the text as possible, or within the Materials and methods section. Journal style for time units are: s, min, h, days, weeks, months, years. Use ‘L’ for litre not ‘l’ to avoid confusion with ‘one’. Use the negative index for units, e.g. number of insects g-1dry wt (also note there is no period for wt). Probability values should be denoted as P.
Mathematical expressions should be carefully represented. Wherever possible, mathematical equations and symbols should be typed in-line by keyboard entry (using Symbol font for Greek characters, and superscript options where applicable). Make sure that there is no confusion between similar characters like l (‘ell’) and 1 (‘one’). Ensure that expressions are spaced as they should appear. If there are several equations they should be identified by an equation number (i.e. ‘eqn 1’ after the equation, and cited in the text as ‘equation 1’).
Numbers from one to nine should be spelled out except when used with units, e.g. two eyes but 10 stomata; 5 °C, 3 years and 5 kg. Do not use excessive numbers of digits when writing a decimal number to represent the mean of a set of measurements. The level of significance implied by numbers based on experimental measurements should reflect, and not exceed, their precision; only rarely can more than 3 figures be justified.