Author contributions
S.S. and W.X. proposed the project. S.S., W.X., and L.B. designed the experiments. L.B., L.H., Y.J., Y.L., Z.Q., C.H., and G.S. performed the experiments. L.B. and S.S. analyzed the data. L.B. wrote the manuscript, with the participation of S.S. and W.Y.