Acknowledgements
This research was supported by the International Research & Development Program of the National Research Foundation of Korea (NRF), funded by the Ministry of Science, ICT & Future Planning (Grant No: 2011-0030500), and by the National Institute of Ecology (Grant No: NIE-C-2020-22) and an Ewha Global Top5 Grant of Ewha Womans University. G. Jeong was supported by the National Research Foundation of Korea (NRF) (Grant No. C00027). Special thanks are given to Mr. Y. Jang, Dr. K. Kobayashi, W. Suehiro, N. Mizumoto, and Y. Nambaat (Kyoto University) and to Dr. M. Okamoto and Dr. S. Mihkeyev (OIST) for commenting on the results and providing Japanese specimens. We thank Dr. Elizabeth Kern (Ewha Womans University) for reviewing the manuscript.