ACKNOWLEDGEMENTS

I thank Da-Yong Zhang, Yanjie Liu, Nianxun Xi, Oliver Bossdorf, and Wenjing Hong for their comments on the manuscript. I thank Xiuxiu Wang and Haidong Qu for their assistance. This study is supported by the National Natural Science Foundation of China (grant No. 31600291 to Y-YZ) and the Fundamental Research Funds for the Central Universities in China (grant No. 20720170074 to Y-YZ).