Author contributions
YZ, WLY, and XLX conceived and designed the research. YZ performed the research. YYZ, DZ, XLT, XH, WHD and ZL participated in the experiments. CS contributed analytic tools. YZ, YYZ, DZ, XLT, and ZL analyzed the experimental data. YZ wrote the manuscript. YYZ, DZ, XLT, and ZL contributed to writing the manuscript. All authors discussed the results and approved the final manuscript.