Author contributions
H.A. and V.A. conducted and analysed the experiments and compiled the data; V.A. and H.A. wrote the manuscript; G.T.S.B. conceived the project, developed the ideas and edited the manuscript; G.V., G.B., X.v.O., W.V.d.E, Y.G and H.As. designed and supervised experiments; D.V. contributed to data analysis; all authors read and approved the manuscript.

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