Acknowledgements
We wish to thank Sierra Oxborrow for assistance with lab work for this project, and Dr. Sarah J. Hart for her feedback on the manuscript. We would also like to acknowledge Veronica Magnusson and the Cedar City Ranger District of the Dixie National Forest for continued support of spruce research on the Markagunt Plateau. This project was made possible through funding from the TW Daniel Endowment, and Utah Agricultural Experiment Station awards 1304 and 1305.