References
Altschul SF, Gish W, Miller W, Myers EW, Lipman DJ. Basic local
alignment search tool. Journal of Molecular Biology. 1990: 215:
403–410.
Beaulieu JM, Moles AT, Leitch IJ, Bennett MD, Dickie JB, Knight CA, et
al. Correlated evolution of genome size and seed mass. 2005;422–37.
Booy G, Raamsdonk LWD Van. Variation in the enzyme esterase within and
between Tulipa species ; usefulness for the analysis of genetic
relationships at different taxonomical levels. Biochem Syst Ecol.
1998;26(2):199-224
Bornnullejr J. Über eine neue Tulpe der Flora Mazedoniens. Feddes
Repert. 1923; 19:199—200.
Christenhusz MJM, Govaerts R, David JC, Hall T, Borland K, Roberts PS,
et al. Tiptoe through the tulips - cultural history, molecular
phylogenetics and classification of Tulipa (Liliaceae). Bot J
Linn Soc. 2013; 172(3):280–328.
Eker I Il, Babaç MTI, Koyuncu M. Revision of the genus Tulipa L.
(Liliaceae) in Turkey. Vol. 157, Phytotaxa. 2014. 1–112 p.
Fay MF, Chase MW, Rønsted N, Devey DS, Pillon… Y. Phylogenetics
of Liliales: summarized evidence from combined analyses of five plastid
and one mitochondrial loci. Aliso [Internet]. 2006;22(November
2015):559–65.
Govaerts R. World Checklist of Selected Plant Families: Royal Botanic
Gardens, Kew [Internet]. Access. 2019 [cited 2019 Aug 13].
Available from: http://wcsp.science.kew.org/qsearch.do
Govaerts, R., Dransfield, J., Zona, S., Hodel, D.R. & Henderson, A.
World Checklist of Arecaceae. Facilitated by the Royal Botanic Gardens,
Kew. Published on the Internet; http://wcsp.science.kew.org/ Retrieved
13 August 2019.
Hayek A. Prodromus Florae peninsulae Balcanicae, Band 3
(Monocotyledoneae). Repertorium specierum Novarum Regni Vegetabilis,
1933; 30:472
Jakob SS, Meister A, Blattner FR. The Considerable Genome Size Variation
of Hordeum Species ( Poaceae ) Is Linked to Phylogeny, Life Form ,
Ecology , and Speciation Rates. Mol Biol Evol. 2004; 21 (5), 860-9
Kress WJ. Plant DNA barcodes: Applications today and in the future. J
Syst Evol. 2017; 55(4):291–307.
Kumar S., Stecher G., Li M., Knyaz C., and Tamura K. MEGA X: Molecular
Evolutionary Genetics Analysis across computing platforms. Molecular
Biology and Evolution. 2018. 35:1547-1549.
Mayer Ernest, Micevski K. Zur Taxonomie und Chorologie von Tulipa
scardica Bornm. Feddes Repertorium. 1970; 591–8.
Millaku F, Elezaj I. Tulipa luanica ( Liliaceae ), a New Species
from Southern Kosovo . Ann Bot Fenn. 2015;52(5–6):315–20.
Millaku F, Elezaj ISA, Berisha N. Sympatric area and ecology of someTulipa species in the west Balkan Peninsula. Thaiszia J Bot.
2018;28(1):35–47.
Obermayer R, Greilhuber J. Does genome size in Dasypyrum villosum vary
with fruit colour? Hredity (Edinb), 2005:91–5.
Osmani M. Disa adaptime citogjenetike dhe biokimike të tre llojeve të
gjinisë Tulipa (T. albanica, T. kosovarica dhe T.
luanica ) në habitatet e tyre natyrore. PhD dissertation. University of
Prishtina; 2018.
Pavlovic, Z. Karakteristicni elementi serpentinske flore Srbije. Glasn.
Prir. Muz. Beogradu, 1962; 8: 3-20.
Peterson A, Levichev IG, Peterson J. Systematics of Gagea andLloydia (Liliaceae) and infrageneric classification of Gagea
based on molecular and morphological data. Mol Phylogenet Evol. 2008;
46(2):446–65.
Raamsdonk LWD Van, Vries T DE. Species relationships and taxonomy inTulipa subg . Tulipa ( Liliaceae ). 1995;195:13–44.
Sang T, Crawford DJ, Stuessy TF. Chloroplast DNA Phylogeny, Reticulate
Evolution, and Biogeography of Paeonia (Paeoniaceae). Am J Bot.
1997; 84 (9),1120–36.
Shuka L, Tan K, Siljak-Yakovlev S. Tulipa albanica (Liliaceae), a new
species from northeastern Albania. Phytotaxa. 2010;10(1):17.
Shuka Lulëzim, Tan Kit KE. Tulipa kosovarica (Liliaceae), a new
species of tulip from Kosovo. Phytotaxa. 2012; 9:1–9.
Swofford, D. L. PAUP*. Phylogenetic Analysis Using Parsimony (*and Other
Methods). 2002. Version 4. Sinauer Associates, Sunderland,
Massachusetts.
Taberlet P., Gielly L., Pautou G., Bouvet J. Universal primers for
amplification of three non-coding regions of chloroplast DNA. Plant
Molecular Biology 1991; 17:1105–1109.
Tatic B, Krivosej Z. Tulipa serbica (Liliaceae), a new species
from Serbia. Bocconea. 1997;5(2):733–6.
The Plant List. Choice Rev Online. 2013; [cited 2019 Aug 15].
Available from: http://www.theplantlist.org/.
Turktas M, Metin ÖK, Baştuǧ B, Ertuǧrul F, Saraç YI, Kaya E. Molecular
phylogenetic analysis of Tulipa (Liliaceae) based on noncoding
plastid and nuclear DNA sequences with an emphasis on Turkey. Bot J Linn
Soc. 2013; 172(3):270–9.
Tutin TG, Heywood VH, Burges NA, Valentine DH. Flora Europaea. Flora
Europaea. 1980. 381–382 p.
Veldkamp JF, Zonneveld BJM. The infrageneric nomenclature ofTulipa (Liliaceae). Plant Syst Evol. 2012; 298(1):87–92.
White T. J., Bruns T. D., Lee S. B., Taylor J. W. 1990. Amplification
and direct sequencing of fungal ribosomal RNA genes for phylogenetics.In M. A. Innis, D. H. Gelfand, J. J. Sninsky, and T. J. White
[eds.], PCR protocols: A guide to methods and applications,
315–322. Academic Press, San Diego, California, USA.
Yanagisawa R, Kuhara T, Nishikawa T, Sochacki D, Marasek-Ciolakowska A,
Okazaki K. Phylogenetic analysis of wild and garden tulips using
sequences of chloroplast DNA. Acta Hortic. 2012; 953:103–10.
Zonneveld BJM, de Groot JJ. Tulipa kolbintsevii Zonn., a new
species from Eastern Kazakhstan. Plant Syst Evol. 2012; 298(7):1293–6.
Zonneveld BJM. The systematic value of nuclear genome size for “ all ”
species of Tulipa L . ( Liliaceae ). Plant Syst Evol. 2009;
281(1):217-245
Zonneveld BJM. The systematic value of nuclear genome size for “all”
species of Tulipa L. (Liliaceae). Plant Syst Evol. 2009;
281(1–2):217–45.
Table 1 . Basic characteristics of the collection sites, voucher
information, and GenBank accession numbers of the Tulipa samples
used for this study