Environmental variables Standardized regression coefficient ± SE
T
F
P
Density
Intercept -109.00 ± 50.18 -2.172 0.035*
Elevation 0.063 ± 0.023 2.768 7.664 0.008**
Soil total nitrogen -11.71 ± 4.54 -2.583 6.671 0.012*
Soil available phosphorus 0.442 ± 0.188 2.356 5.551 0.022*
Adult density 0.007 ± 0.003 2.268 5.142 0.027*
Species richness
Intercept -12.43 ± 25.69 -0.484 0.631
Soil total nitrogen -6.815 ± 2.405 -2.834 8.030 0.006**
Elevation 0.026 ± 0.012 2.249 5.060 0.029*
Canopy density 25.03 ± 11.55 2.167 4.696 0.034*