Acknowledgements

This project was funded through a British Ecological Society Research grant (4849/5889) and Severo Ochoa grants (MicroP_SO_14 and MicroProy_SO_15) from the EBD-CSIC. OR was funded through a Natural Environment Research Council Independent Research Fellowship (NE/M018660/1) and University of Stirling Impact Fellowship. We are grateful to Neguse Engda, Ziyad Jemal and Dessiew Gelaw for help with fieldwork, and Laura Kubasiewicz and Juan Luís García-Mudarra of the Molecular Ecology lab LEM-EBD for carrying out the lab work. Research and export permits were granted by the Ethiopian Wildlife Conservation Authority (EWCA) with the help of Prof Afework Bekele (Addis Ababa University). We thank the wardens and experts of Bale Mountains National Park, Arsi Mountains National Park, Simien Mountains National Park, Aboy Gara Community Conservation and Guassa Community Based Conservation Area. The following people and organisations facilitated the fieldwork in Ethiopia: Isabel Cantero (CSIC and Addis Ababa University), Claudio Sillero and the Ethiopian Wolf Conservation project, Karen Laurenson and Frankfurt Zoological Society-Ethiopia (FZS-ETH), Spanish Embassy in Ethiopia, Guy and Yvonne Levene of Bale Mountain Lodge, Guassa Lodge, and Manner Tour Car Rental and car drivers.