Acknowledgements
We thank the Smithsonian Tropical Research Institute for facilitating our fieldwork in Panama. Our field survey at Lambir Hills National Park was conducted in accordance with the Memorandums of Understanding signed between the Sarawak Forest Department (SFD, Kuching, Malaysia) and the Japan Research Consortium for Tropical Forests in Sarawak (JRCTS, Sendai, Japan) in December 2012. Thanks to Mohd. Shahbudin Sabki, Engkamat Lading, and Mohamad bin Kohdi, Paulus Meleng of SFD for help in obtaining research permission at the Lambir Hills National Park. We thank Tohru Nakashizuka (Tohoku University, Sendai, Japan) and Tomoaki Ichie (Kochi University, Kochi, Japan) for their support. Diaspidid sample processing was by Katelyn Mullen, Ryan McCarthy, Mitchel Logan, Michael Fizdale, Kyara Romeu, Shannon Trujillo, Hannah Shapiro, and Anna Whitfield. DAP’s dissertation committee (Michael Hood, Adam Porter and Laura A. Katz) and anonymous reviewers provided helpful feedback. JW’s participation was supported by the China Scholarship Council. This work was supported by the National Science Foundation (DEB-1258001; DEB-1744552) and by Grants-in-Aid (no. 21255004 to TI) from the Japanese Ministry of Education, Science and Culture.