Author contributions
JP, GH, MB and MAS assisted in the conception and design of the study. MAS, EZ, SR and JD assisted in assembling BOLD datasets and providing DNA extracts for laboratory experiments. Field and Laboratory work was undertaken by JP and PT. SRA work was undertaken by HD and SS. Analysis and interpretation of the data were undertaken by JP, PT, HD, GH, MB and SS, as well as drafting of the manuscript. All authors assisted in critical revision of the manuscript.