Author contributions
JP, GH, MB and MAS assisted in the conception and design of the study.
MAS, EZ, SR and JD assisted in assembling BOLD datasets and providing
DNA extracts for laboratory experiments. Field and Laboratory work was
undertaken by JP and PT. SRA work was undertaken by HD and SS. Analysis
and interpretation of the data were undertaken by JP, PT, HD, GH, MB and
SS, as well as drafting of the manuscript. All authors assisted in
critical revision of the manuscript.