Author contributions
H.D. and X.Y.C. designed, conducted, and analyzed the experiments with oversight by Y.H.Z.. Y.T.L. and X.H. performed forced swim test. H.L.Z and Y.Q. performed sucrose preference test. N.K. performed novelty-suppressed feeding test. H.D., S.Y.C. and Y.H.Z. wrote the paper. All authors reviewed and approved the final manuscript.