ACKNOWLEDGMENTS
This work was funded by the National Key Research and Development Program of China (grant number 2017YFD0600105), National Natural Science Foundation of China (grant numbers U1905201 and 31601905), Science and Technology Program of Fujian Province (grant number 2018N5002), Forestry Science Research Project of Fujian Forestry Department (grant number Minlinke [2017] 03).We are very grateful to Jianghua Sun (State Key Laboratory of Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Acade my of Sciences, Beijing, China) for help and support in this study.We warmly thank Key Laboratory of Integrated Pest Management in Ecological Forests, Fujian Province University for providing experimental environment, as well as thank Shunchang County Forestry Bureau and Xiapu County Forestry Bureau for providing research areas and helping to complete the collection of all samples.