Acknowledgements
Data for this study were collected by the former SYSU-Alberta Joint Lab for Biodiversity Conservation. This work was supported by the National Natural Science Foundation of China (Grant NO. 31901107) and Natural Science Foundation of Guangdong Province (Grant NO. 2018A030310384). DY gratefully acknowledges financial support from China Scholarship Council and Mitacs Globalink Early Career Fellowship-China. MWC and FH were both supported by the Natural Sciences and Engineering Research Council of Canada.
Author contributions: DY, MWC and FH designed the study. DY led the analyses. DY, MWC and FH wrote the manuscript with contribution from all authors.
Conflict of Interest: The authors declare no conflict of interest.