References
Addison, P., & Samways, M. (2006). Surrogate habitats demonstrate the
invasion potential of the African pugnacious ant. Biodiversity and
Conservation, 15 (1), 411-428. doi:10.1007/s10531-005-5399-4
Addison, P., & Samways, M. J. (2000). A survey of ants (Hymenoptera:
Formicidae) that forage in vineyards in theWestern Cape Province, South
Africa. African Entomology, 8 , 251–260.
Ahuatzin, D. A., Corro, E. J., Jaimes, A. A., Valenzuela Gonzalez, J.
E., Feitosa, R. M., Ribeiro, M. C., . . . Dattilo, W. (2019). Forest
cover drives leaf litter ant diversity in primary rainforest remnants
within human-modified tropical landscapes. Biodiversity and
Conservation, 28 (5), 1091-1107. doi:10.1007/s10531-019-01712-z
Andersen, A. N. (2019). Responses of ant communities to disturbance:
Five principles for understanding the disturbance dynamics of a globally
dominant faunal group. Journal of Animal Ecology, 88 (3), 350-362.
doi:10.1111/1365-2656.12907
Archer, S. R., Andersen, E. M., Predick, K. I., Schwinning, S., Steidl,
R. J., & Woods, S. R. (2017). Woody Plant Encroachment: Causes and
Consequences. In D. D. Briske (Ed.), Rangeland Systems: Processes,
Management and Challenges (pp. 25-84).
Baker, M. E., & King, R. S. (2010). A new method for detecting and
interpreting biodiversity and ecological community thresholds.Methods in Ecology and Evolution, 1 (1), 25-37.
doi:10.1111/j.2041-210X.2009.00007.x
Bestelmeyer, B. T. (2005). Does desertification diminish biodiversity?
Enhancement of ant diversity by shrub invasion in south-western USA.Diversity and Distributions, 11 (1), 45-55.
doi:10.1111/j.1366-9516.2005.00122.x
Blaum, N., Seymour, C., Rossmanith, E., Schwager, M., & Jeltsch, F.
(2009). Changes in arthropod diversity along a land use driven gradient
of shrub cover in savanna rangelands: identification of suitable
indicators. Biodiversity and Conservation, 18 (5), 1187-1199.
doi:10.1007/s10531-008-9498-x
Bond, W. J. (2019). Open Ecosystems: Ecology and Evolution beyond
the Forest Edge . New York: Oxford Univ Press.
Botes, A., McGeoch, M. A., Robertson, H. G., Van Niekerk, A., Davids, H.
P., & Chown, S. L. (2006). Ants, altitude and change in the northern
Cape Floristic Region. Journal of Biogeography, 33 , 71-90.
Boundja, R. P., & Midgley, J. J. (2010). Patterns of elephant impact on
woody plants in the Hluhluwe-Imfolozi park, Kwazulu-Natal, South Africa.African Journal of Ecology, 48 (1), 206-214.
doi:10.1111/j.1365-2028.2009.01104.x
Bownes, A., Moore, S. D., & Villet, M. H. (2014). My enemy’s enemies:
recruiting hemipteran-tending generalist ants for biological control in
citrus orchards by spatial partitioning of foraging webs. African
Entomology, 22 (3), 519-529. doi:10.4001/003.022.0303
Campos, R. I., Soares, J. P., Martins, R. P., & Ribeiro, S. P. (2006).
Effect of habitat structure on ant assemblages (Hymenoptera :
Formicidae) associated to two pioneer tree species. Sociobiology,
47 (3), 721-737.
Charles-Dominique, T., Staver, A. C., Midgley, G. F., & Bond, W. J.
(2015). Functional differentiation of biomes in an African
savanna/forest mosaic. South African Journal of Botany, 101 ,
82-90. doi:10.1016/j.sajb.2015.05.005
Costas, N., Pardo, I., Mendez-Fernandez, L., Martinez-Madrid, M., &
Rodriguez, P. (2018). Sensitivity of macroinvertebrate indicator taxa to
metal gradients in mining areas in Northern Spain. Ecological
Indicators, 93 , 207-218. doi:10.1016/j.ecolind.2018.04.059
Criado, M. G., Myers-Smith, I. H., Bjorkman, A. D., Lehmann, C. E. R.,
& Stevens, N. (2020). Woody plant encroachment intensifies under
climate change across tundra and savanna biomes. Global Ecology
and Biogeography, 29 (5), 925-943. doi:10.1111/geb.13072
Dean, W. R. J. (1992). Temperatures determining activity patterns of
some ant species in the Southern Karoo, South Africa. Journal of
the Entomological Society of Southern Africa, 55 (1), 149-156.
Del Toro, I., Ribbons, R. R., & Pelini, S. L. (2012). The little things
that run the world revisited: a review of ant-mediated ecosystem
services and disservices (Hymenoptera: Formicidae). Myrmecological
News, 17 , 133-146.
Eldridge, D. J., Bowker, M. A., Maestre, F. T., Roger, E., Reynolds, J.
F., & Whitford, W. G. (2011). Impacts of shrub encroachment on
ecosystem structure and functioning: towards a global synthesis.Ecology Letters, 14 (7), 709-722.
doi:10.1111/j.1461-0248.2011.01630.x
Fisher, B. L., & Bolton, B. (2016). Ants of Africa and
Madagascar: a guide to the genera . United States: University of
California Press.
Gray, E. F., & Bond, W. J. (2013). Will woody plant encroachment impact
the visitor experience and economy of conservation areas? Koedoe,
55 (1), 9. doi:10.4102/koedoe.v55i1.1106
Greve, M. E., Blaha, S., Teuber, J., Rothmaier, M., & Feldhaar, H.
(2019). The effect of ground surface rugosity on ant running speed is
species-specific rather than size dependent. Insectes Sociaux,
66 (3), 355-364. doi:10.1007/s00040-019-00694-z
Guilherme, D. R., Souza, J. L. P., Franklin, E., Pequeno, P., das
Chagas, A. C., & Baccaro, F. B. (2019). Can environmental complexity
predict functional trait composition of ground-dwelling ant assemblages?
A test across the Amazon Basin. Acta Oecologica-International
Journal of Ecology, 99 , 27. doi:10.1016/j.actao.2019.05.004
Hering, R., Hauptfleisch, M., Geissler, K., Marquart, A., Schoenen, M.,
& Blaum, N. (2019). Shrub encroachment is not always land degradation:
Insights from ground-dwelling beetle species niches along a shrub cover
gradient in a semi-arid Namibian savanna. Land Degradation &
Development, 30 (1), 14-24. doi:10.1002/ldr.3197
Hethcoat, M. G., King, B. J., Castiblanco, F. F., Ortiz-Sepulveda, C.
M., Achiardi, F. C. P., Edwards, F. A., . . . Edwards, D. P. (2019). The
impact of secondary forest regeneration on ground-dwelling ant
communities in the Tropical Andes. Oecologia, 191 (2), 475-482.
doi:10.1007/s00442-019-04497-8
Hölldobler, B., & Wilson, E. O. (1990). The ants . Massachusetts:
Harvard University Press.
Hui, F. K. (2016). boral–Bayesian ordination and regression analysis of
multivariate abundance data in R. Methods in Ecology and
Evolution, 7 (6), 744-750.
Jost, L. (2006). Entropy and diversity. Oikos, 113 (2), 363-375.
Keiser, C. N., Wright, C. M., & Pruitt, J. N. (2015). Warring arthropod
societies: Social spider colonies can delay annihilation by predatory
ants via reduced apparency and increased group size. Behavioural
Processes, 119 , 14-21. doi:10.1016/j.beproc.2015.07.005
Kettenbach, J. A., Miller-Struttmann, N., Moffett, Z., & Galen, C.
(2017). How shrub encroachment under climate change could threaten
pollination services for alpine wildflowers: A case study using the
alpine skypilot, Polemonium viscosum. Ecology and Evolution,
7 (17), 6963-6971. doi:10.1002/ece3.3272
Koch, B., Edwards, P. J., Blanckenhorn, W. U., Walter, T., & Hofer, G.
(2015). Shrub encroachment affects the diversity of plants, butterflies,
and grasshoppers on two Swiss subalpine pastures. Arctic Antarctic
and Alpine Research, 47 (2), 345-357. doi:10.1657/aaar0013-093
Kwok, A. B. C., & Eldridge, D. J. (2016). The influence of shrub
species and fine-scale plant density on arthropods in a semiarid
shrubland. Rangeland Journal, 38 (4), 381-389. doi:10.1071/rj15019
Lara-Romero, C., Garcia, C., Morente-Lopez, J., & Iriondo, J. M.
(2016). Direct and indirect effects of shrub encroachment on alpine
grasslands mediated by plant-flower visitor interactions.Functional Ecology, 30 (9), 1521-1530. doi:10.1111/1365-2435.12637
Lassau, S. A., & Hochuli, D. F. (2004). Effects of habitat complexity
on ant assemblages. Ecography, 27 (2), 157-164.
doi:10.1111/j.0906-7590.2004.03675.x
Mauda, E. V., Joseph, G. S., Seymour, C. L., Munyai, T. C., & Foord, S.
H. (2018). Changes in landuse alter ant diversity, assemblage
composition and dominant functional groups in African savannas.Biodiversity and Conservation, 27 (4), 947-965.
doi:10.1007/s10531-017-1474-x
Mitchell, P. L. (2000). Leaf-footed bugs (Coreidae). In C. W. Schaefer
& A. R. Panizzi (Eds.), Heteroptera of economic importance .
Florida: CRC Press.
Munyai, T. C., & Foord, S. H. (2012). Ants on a mountain: spatial,
environmental and habitat associations along an altitudinal transect in
a centre of endemism. Journal of Insect Conservation, 16 ,
677-695. doi:10.1007/s10841-011-9449-9
Munyai, T. C., & Foord, S. H. (2015). An inventory of epigeal ants of
the western Soutpansberg Mountain Range, South Africa. koedoe,
57 (1), 1-12.
Nakagawa, S., & Schielzeth, H. (2013). A general and simple method for
obtaining R2 from generalized linear mixed‐effects models. Methods
in Ecology and Evolution, 4 (2), 133-142.
Nooten, S. S., Schultheiss, P., Rowe, R. C., Facey, S. L., & Cook, J.
M. (2019). Habitat complexity affects functional traits and diversity of
ant assemblages in urban green spaces (Hymenoptera: Formicidae).Myrmecological News, 29 , 67-77.
doi:10.25849/myrmecol.news_029:067
O’Connor, T. G., Puttick, J. R., & Hoffman, M. T. (2014). Bush
encroachment in southern Africa: changes and causes. African
Journal of Range & Forage Science, 31 (2), 67-88.
doi:10.2989/10220119.2014.939996
Osborne, C. P., Charles-Dominique, T., Stevens, N., Bond, W. J.,
Midgley, G., & Lehmann, C. E. R. (2018). Human impacts in African
savannas are mediated by plant functional traits. New Phytologist,
220 (1), 10-24. doi:10.1111/nph.15236
Pacheco, R., & Vasconcelos, H. (2012). Habitat diversity enhances ant
diversity in a naturally heterogeneous Brazilian landscape.Biodiversity and Conservation, 21 (3), 797-809.
doi:10.1007/s10531-011-0221-y
Parr, C. L. (2008). Dominant ants can control assemblage species
richness in a South African savanna. Journal of Animal Ecology,
77 (6), 1191-1198. doi:10.1111/j.1365-2656.2008.01450.x
Parr, C. L., Gray, E. F., & Bond, W. J. (2012). Cascading biodiversity
and functional consequences of a global change-induced biome switch.Diversity and Distributions, 18 (5), 493-503.
doi:10.1111/j.1472-4642.2012.00882.x
Parui, A. K., Chatterjee, S., & Basu, P. (2015). Habitat
characteristics shaping ant species assemblages in a mixed deciduous
forest in Eastern India. Journal of Tropical Ecology, 31 ,
267-280. doi:10.1017/s0266467415000036
Pausas, J. G., & Bond, W. J. (2020). Alternative Biome States in
Terrestrial Ecosystems. Trends in Plant Science, 25 (3), 250-263.
doi:10.1016/j.tplants.2019.11.003
Price, J. N., & Morgan, J. W. (2008). Woody plant encroachment reduces
species richness of herb-rich woodlands in southern Australia.Austral Ecology, 33 (3), 278-289.
doi:10.1111/j.1442-9993.2007.01815.x
Prins, A. J., Robertson, H. G., & Prins, A. (1990). Pest ants in urban
and agricultural areas of southern Africa. In R. Van der Meer, K. Jaffe,
& A. Cedeno (Eds.), Applied myrmecology: a world perspective .
Boulder: Westview Press.
Radnan, G. N., Gibb, H., & Eldridge, D. J. (2018). Soil surface
complexity has a larger effect on food exploitation by ants than a
change from grassland to shrubland. Ecological Entomology, 43 (3),
379-388. doi:10.1111/een.12510
Rios-Casanova, L., Valiente-Banuet, A., & Rico-Gray, V. (2006). Ant
diversity and its relationship with vegetation and soil factors in an
alluvial fan of the Tehuacan Valley, Mexico. Acta
Oecologica-International Journal of Ecology, 29 (3), 316-323.
doi:10.1016/j.actao.2005.12.001
Samways, M. (1983). Community structure of ants (Hymenoptera:
Formicidae) in a series of habitats associated with citrus.Journal of Applied Ecology, 20 , 833-847.
Samways, M. (1990). Species temporal variability: epigaeic ant
assemblages and management for abundance and scarcity. Oecologia,
84 , 482-490.
Sirami, C., & Monadjem, A. (2012). Changes in bird communities in
Swaziland savannas between 1998 and 2008 owing to shrub encroachment.Diversity and Distributions, 18 (4), 390-400.
doi:10.1111/j.1472-4642.2011.00810.x
Stevens, N., Lehmann, C. E., Murphy, B. P., & Durigan, G. (2017).
Savanna woody encroachment is widespread across three continents.Global change biology, 23 (1), 235-244.
Vermeij, G. J. (1995). Species-diversity in space and time - Rosenzweig,
ML. Science, 270 (5235), 503-504.
Wang, Y., Naumann, U., Wright, S. T., & Warton, D. I. (2012). mvabund-
an R package for model-based analysis of multivariate abundance data.Methods in Ecology and Evolution, 3 (3), 471-474.
doi:10.1111/j.2041-210X.2012.00190.x
Warton, D. I., Thibaut, L., & Wang, Y. A. (2017). The PIT-trap-A
”model-free” bootstrap procedure for inference about regression models
with discrete, multivariate responses. PLoS ONE, 12 (7).
doi:10.1371/journal.pone.0181790
Warton, D. I., Wright, S. T., & Wang, Y. (2012). Distance-based
multivariate analyses confound location and dispersion effects.Methods in Ecology and Evolution, 3 (1), 89-101.
doi:10.1111/j.2041-210X.2011.00127.x
Wills, B. D., & Landis, D. A. (2018). The role of ants in north
temperate grasslands: a review. Oecologia, 186 (2), 323-338.
doi:10.1007/s00442-017-4007-0
Yusah, K. M., & Foster, W. A. (2016). Tree size and habitat complexity
affect ant communities (Hymenoptera: Formicidae) in the high canopy of
Bornean rain forest. Myrmecological News, 23 , 15-23.
Table 1: Best models and the marginal and conditional variation
explained based on Generalized Linear Mixed Effects Models (GLMM) of
species richness, Shannon and Simpsons diversity. Shannon and Simpson
indices are represented by there effective numebers