Acknowledgements
This research was supported by grants from the National Research Foundation of Korea (2012K1A2B1A03000496) funded by the Government of the Republic of Korea (MSIP), and the Research Institute of Basic Sciences, Seoul National University (to B.W.), and from the National Natural Science Foundation of China (31211140342) (to P.L.), SINO-BON (to J.P.J.). We thank members of the Min and Waldman labs for help with fieldwork.