Acknowledgements
Funding was provided by the National Science Foundation (HRD-1826490), the US Department of Education HSI-STEM grant, and a California State University Dominguez Hills RSCA award, and a Graduate Student Fellowship from the Organization for Tropical Studies (OTS). We would like to thank the staff of the OTS, particularly Danilo Brenes and Bernal Matarrita, for their immeasurable support. We thank E. Kuprewicz for advice on analysis, R.F. de CaƱa and J.E. Watkins for field supplies, and the Urban Nature Research Center of the Natural History Museum for feedback on the manuscript, including K. Bell, B. Brown, A. Clause, M. Connolly, G. Pauly, A. Schulz, and J. Vendetti. This work was conducted under a research permit issued by the Costa Rican Ministry of Energy and Environment.