Acknowledgements

We thank Ryota Aoyagi, Keito Kobayashi, Susan Cordell, Elizabeth Stacy, Rebecca Ostertag, Amanda Uowolo, Tomoko Sakishima, Gary Sur, and Joshua Nagle for their supports on our fieldwork. We also thank the Department of Land and Natural Resources (DOFAW) for permission to take plant samples. This study was partly supported by grants from JSPS KAKENHI #JP17J04193 to GA and #15KK0255 to YO, and Internship Subsidy from CoHHO (the Connectivity of Hills, Humans, and Oceans) Educational Program to GA.