Acknowledgements
This study was supported by Natural Science Foundation of Shandong Province of China (ZR2019QC002), Key Laboratory of Healthy Freshwater Aquaculture, Ministry of Agriculture and Rural affairs; Key Laboratory of Freshwater Aquaculture genetic and breeding of Zhejiang Province; Zhejiang Institute of Freshwater Fisheries, Huzhou (ZJK202003), Agricultural Science and Technology independent innovation project (CX(19)2034), Creation Project of Major New Species of Agriculture of Jiangsu Province(PZCZ201742) and Qingchuang science and technology support program of Shandong provincial college.