Author Contributions:
HF Cooper collected trait data, performed genetic and statistical analyses, and wrote much of the manuscript. LV Andrews helped with RADseq processing. RJ Best contributed essential analysis of our results and helped write the manuscript. KR Grady assisted with field collections, and along with GJ Allan, CA Gehring, KR Hultine, and TG Whitham, designed the common garden experiment. All authors provided critical edits to the manuscript and figures.