References
Anderson, A. M., Duijns, S., Smith, P. A., Friis, C. and Nol, E. 2019.
Migration distance and body condition influence shorebird migration
strategies and stopover decisions during southbound migration. – Front.
Ecol. Evol. 7: 251.
Bentzen, R., Dondua, A., Porter, R., Robard, M. D. and Solovyeva, D.V.
2016. Large-scale movements of Dunlin breeding in Chukotka, Russia,
during the non-breeding period. – Wader Study 132: 1–14.
Boulet, M. and Norris, D. R. 2006. Introduction: The Past and Present of
Migratory Connectivity. – Ornithol. Monogr. 61: 1–13.
Chen, K., Yang, X. and Lu, Y. 2015. Vital stopover of shorebirds
migration on the East Asian-Australasian flyway: Wetlands of yellow sea
and Bohai Sea. – Wetl. Sci. 13: 1–6.
Choi. C., Hua, N., Persson, C., Chiang, C. and Ma, Z. 2010. Age-related
plumage differences of Dunlins along the East Asian-Australasian Flyway.
– J. Field Ornithol. 81: 99–111.
Darriba, D., Taboada, G.L., Doallo, R. and Posada, D. 2012. jModelTest
2: more models, new heuristics and parallel computing. – Nat. Methods
9: 772–772.
Dunn, E. H. 2001. Mass change during migration stopover: a comparison of
species groups and sites. – J. Field Ornithol. 72: 419–432.
Gates, H.R., Yezerinac, S., Powell, A. N., Tomkovich, P. S., Valchuk, O.
P. and Lanctot, R. B. 2013. Differentiation of subspecies and sexes of
Beringian Dunlins using morphometric measures. – J. Field Ornithol. 84:
389–402.
Gill, R. E., Handel, C. M. and Ruthrauff, D. R. 2013. Intercontinental
migratory connectivity and population structuring in Dunlins
(Calidris alpina ) from western Alaska. – Condor 115: 525–534.
Gill, R. E., Tibbitts, T. L., Douglas, D. C., Handel, C. M., Mulcahy, D.
M., Gottschalck, J. C., Warnock, N., Mccaffery, B. J., Battley, P. F.
and Piersma, T. 2009. Extreme endurance flights by landbirds crossing
the Pacific Ocean: ecological corridor rather than barrier? – P. Roy.
Soc. B-Biol Sci. 276: 447–457.
Grattarola, A., Spina, F. and Pilastro, A. 1999. Spring migration of the
Garden Warbler (Sylvia borin ) across the Mediterranean Sea. – J.
Ornithol. 140: 419–430.
Greenwood, J. G. 1984. Migration of dunlin calidris alpina: A worldwide
overview. – Ring. Migr. 5: 35–39.
Greenwood,
J. G. 1986. Geographical variation and taxonomy of the DunlinCalidris alpine . – Bull. Brit. Ornithol. Club 106: 284–286.
Griffiths, R., Double, M. C., Orr, K. and Dawson, R. J. G. 1998. A DNA
test to sex most birds. – Mol. Ecol. 7: 1071–1075.
Gromadzka, J. 1989. Breeding and Wintering Areas of Dunlin Migrating
through Southern Baltic. – Ornis Scand. 20: 132–144.
Guindon, S., Dufayard, J. F., Lefort, V., Anisimova, M., Hordijk, W. and
Gascuel, O. 2010. New algorithms and methods to estimate
maximum-likelihood phylogenies: assessing the performance of PhyML 3.0.
– Syst. Biol. 59: 307–321.
Jahn, A. E., Levey, D. J., Cueto, V. R., Ledezma, J. P., Tuero, D. T.,
Fox, J. W. and Masson, D. 2013. Long-Distance Bird Migration within
South America Revealed by Light-Level Geolocators. – Auk 130: 223–229.
Jing-Gong, X., Ma, Z. J., Yang, L. and Chen, J. K. 2009. Progress in
infraspecific taxonomy of dunlin Calidris alpona . – Acta Zootax.
Sin. 34: 546–553.
Johnston,
V., Syroechkovskiy, E., Crockford, N., Lanctot, R. B., Millington, S.,
Clay, R., Donaldson, G., Ekker, M., Gilchrist, G., Black, A. and
Crawford, R. 2015. Arctic Migratory Birds Initiative (AMBI): Workplan
2015-2019. CAFF Strategies Series, 6.
Kumar, S., Stecher, G. and Tamura, K. 2016. MEGA7: Molecular
Evolutionary Genetics Analysis Version 7.0 for Bigger Datasets. – Mol.
Biol. Evol. 33: 1870–1874.
Lagassé, B. J., Lanctot, R. B., Barter, M., Brown, S., Chiang, C. Y.,
Choi, C. Y., Gerasimov Y. N., Kendall, S., Liebezeit. J. R., Maslovsky,
K. S., Matsyna, A. I., Matsyna, E. L., Payer, D. C., Saalfeld, S. T.,
Shigeta, Y., Tiunov, I. M., Tomkovich, P. S., Valchuk, O. P. and Wunder,
M. B. 2020. Dunlin subspecies exhibit regional segregation and high site
fidelity along the East Asian–Australasian Flyway. – Condor 122: 1–5.
Lagassé, B. J., Lanctot, R. B., Brown, S., Dondua, A. G., Kendall, S.,
Latty, C. J., Liebezeit J. R., Loktionov E. Y., Maslovsky, K. S.,
Matsyna, A. I., Matsyna, E. L., McGuire, R. L., Payer, D. C., Saalfeld,
S. T., Slaght, J. C., Solovyeva, D. V., Tomkovich, P. S., Valchuk, O. P.
and Wunder, M. B. 2022. Migratory network reveals unique
spatial-temporal migration dynamics of Dunlin subspecies along the East
Asian-Australasian Flyway. – PLoS One 17: e0270957.
Li, H., Mao, D., Wang, Z., Huang, X., Li, L. and Jia, M. 2022. Invasion
of Spartina alterniflora in the coastal zone of mainland China: Control
achievements from 2015 to 2020 towards the Sustainable Development
Goals. – J. Environ. Manage. 323: 116242.
Librado, P. and Rozas, J. 2009. DnaSP v5: a software for comprehensive
analysis of DNA polymorphism data. – Bioinformatics 25: 1451–1452.
Lisovski, S., Gosbell, K., Hassell, C. and Minton, C. 2016. Tracking the
full annual-cycle of the Great Knot, Calidris tenuirostris, a
long-distance migratory shorebird of the East Asian-Australasian Flyway.
– Wader Study123: 1–12.
Lopes, R. J., Marques, J. C. and Wennerberg, L. 2006. Migratory
connectivity and temporal segregation of dunlin (Calidris alpina )
in Portugal: evidence from morphology, ringing recoveries and mtDNA. –
J.Ornithol. 147: 385–394.
Ma, Y., Choi, C. Y., Thomas, A. and Gibson, L. 2022. Review of
contaminant levels and effects in shorebirds: Knowledge gaps and
conservation priorities. – Ecotox. Environ. Safe. 242: 113868.
Ma, Z., Hua, N., Peng, H., Choi, C., Battley, P. F., Zhou, Q., Chen, Y.,
Ma, Q., Jia, N. and Xue, W. 2013. Differentiating between stopover and
staging sites: functions of the southern and northern Yellow Sea for
long-distance migratory shorebirds. – J. Avian Biol. 44: 504–512.
Ma, Z., Melville, D. S., Liu, J., Chen, Y., Yang, H., Ren, W., Zhang,
Z., Piersma, T. and Li, B. 2014. Rethinking China’s new great wall. –
Science 346: 912–914.
Marthinsen, G., Wennerberg, L. and Lifjeld, J. T. 2007. Phylogeography
and subspecies taxonomy of dunlins (Calidris alpina ) in western
Palearctic analysed by DNA microsatellites and amplified fragment length
polymorphism markers. – Biol. J. Linn. Soc. 92: 713–726.
Melville, D. S., Chen, Y. and Ma, Z. 2016. Shorebirds along the Yellow
Sea coast of China face an uncertain future—a review of threats. –
Emu 116: 100–110.
Miller, M. P., Haig, S. M., Mullins, T. D., Ruan, L., Casler, B.,
Dondua, A., Gates, H. R., Johnson, J. M., Kendall, S. and Tomkovich, P.
S. 2014. Intercontinental genetic structure and gene flow in Dunlin
(Calidris alpina ), a potential vector of avian influenza. –
Evol. Appl. 8: 149–171.
Moore, F. R. and Simons, T. R. 1990. Stopover on a Gulf Coast Barrier
Island by Spring Trans-Gulf Migrants. – Wilson Bull. 102: 487–500.
Mu, T., Tomkovich, P. S., Loktionov, E. Y., Syroechkovskiy, E. E. and
Wilcove, D. S. 2018. Migratory routes of red‐necked phalaropes
Phalaropus lobatus breeding in southern Chukotka revealed by
geolocators. – J. Avian Biol. 49: e01853.
Popovic, I., Toews, D. P., Keever,
C. C., St. Clair, C. T., Barbaree, B. A., Fernández, G. and Rourke, J.
2019. Genotyping-by-sequencing reveals genomic homogeneity among
overwintering Pacific Dunlin (Calidris alpina pacifica )
aggregations along the Pacific coast of North America. – Condor 121:
duz036.
Rheindt, F. E., Székely, T., Edwards, S. V., Lee, P. L., Burke, T.,
Kennerley, P. R., Bakewell, D. N., Alrashidi, M., Kosztolányi, A. and
Weston, M. A. 2011. Conflict between genetic and phenotypic
differentiation: the evolutionary history of a ’lost and rediscovered’
shorebird. – PLoS One 6: e26995.
Ronquist, F., Teslenko, M., Mark, P. V. D., Ayres, D. L., Darling, A.,
Höhna, S., Larget, B., Liu, L., Suchard, M.A. and Huelsenbeck, J. P.
2012. MrBayes 3.2: efficient Bayesian phylogenetic inference and model
choice across a large model space. – Syst. Biol. 61: 539–542.
Sambrook, J. E., Maniatis, T. E. and Fritsch, E. F. 1989. Molecular
Cloning: A Laboratory Manual Cold Spring Harbor Laboratory, Cold Spring
Harbor, NY. 1.
Shumway, N., Saunders, M. I., Nicol, S., Fuller, R. A., Ben‐Moshe, N.,
Iwamura, T., Kim, S. W. Murray, N. J., Watson, J. E. M. and Maron, M.
2022. Exploring the risks and benefits of flexibility in biodiversity
offset location in a case study of migratory shorebirds. – Conserv.
Biol. 37: e14031.
Smith, H. G. and Schantz, T. V. 1996. Adoption or Infanticide: Options
of Replacement Males in the European Starling. – Behav. Ecol.
Sociobiol. 38: 191–197.
Soikkeli, M. 1967. Breeding cycle and population dynamics in the dunlin
(Calidris alpina ). – Ann. Zool. Fenn. 4: 158–198.
Studds, C. E., Kendall, B. E., Murray, N. J., Wilson, H. B., Rogers, D.
I., Clemens, R. S., Gosbell, K., Hassell, C. J., Jessop, R., Melville,
D. S., Milton, D. A., Minton, C. D. T., Possingham, H. P., Riegen, A.
C., Straw, P., Woehler, E. J. and Fuller, R. A. 2017. Rapid population
decline in migratory shorebirds relying on Yellow Sea tidal mudflats as
stopover sites. – Nat. Commun. 8: 14895.
Sutherland, W. J., Alves, J. A., Amano, T., Chang, C. H., Davidson, N.
C., Max Finlayson, C., Gill, J. A., GillJr, R. E., González, P. M.,
Gunnarsson, T. G., Kleijn, D., Spray, C. J., Székely, T. and Thompson,
D. B. A. 2012. A horizon scanning assessment of current and potential
future threats to migratory shorebirds. – Ibis 154: 663–679.
Tomkovich, P. S. 1986. Geographical variability of the Dunlin in the Far
East. – Bull. Mosc. Soc. Nat. 91: 3–15.
Tong, M., Lin, Z., Li, J., Zöckler, C. and Clark, N. 2012. The critical
importance of the Rudong mudflats, Jiangsu Province, China in the annual
cycle of the Spoon-billed Sandpiper Calidris pygmeus . – Ciba
Found. Stud Gr. 119: 74–77.
Verkuil, Y. I., Piersma, T., Jukema, J., Hooijmeijer, J. C. E. W.,
Zwarts, L. and Baker, A. J. 2012. The interplay between habitat
availability and population differentiation: a case study on genetic and
morphological structure in an inland wader (Charadriiformes). – Biol.
J. Linn. Soc. 106: 641–656.
Villesen, P. 2007. FaBox: an online toolbox for fasta sequences. – Mol.
Ecol. Notes 7: 965–968.
Wang, X., Chen, Y., Melville, D. S., Choi, C. Y., Tan, K., Liu, J., Li,
J., Zhang, S. Cao, L. and Ma, Z. 2022. Impacts of habitat loss on
migratory shorebird populations and communities at stopover sites in the
Yellow Sea. – Biol. Conserv. 269: 109547.
Warnock, N. 2010. Stopping vs. staging: the difference between a hop and
a jump. – J. Avian Biol. 41: 621–626.
Webster, M. S., Marra, P. P., Haig, S. M., Bensch, S. and Holmes, R. T.
2002. Links between worlds: unraveling migratory connectivity. – Trends
Ecol. Evol. 17: 76–83.
Weiser, E. L., Lanctot, R. B., Brown, S. C., Gate, H. R., Bentzen, R.
L., Bety, J., Boldenow, M. L., English, W. B., Franks, S. E., Koloski,
L., Kwon, E., Lamarre, J. F., Lank, D. B., Liebezeit, J. R., Mckinnon,
L., Nol, E., Rausch, J., Saalfeld, S. T., Senner, N. R., Ward, D. H.,
Woodard, P. F. and Sandercock, B. K. 2018. Environmental and ecological
conditions at Arctic breeding sites have limited effects on true
survival rates of adult shorebirds. – Auk 135: 29–43.
Wenink, P. W. and Baker, A. J. 1996. Mitochondrial DNA lineages in
composite flocks of migratory and wintering dunlins (Calidris
alpina ). – Med. Phys. 38: 3611–3611.
Wenink, P. W. & Tilanus, M. G. 1996. Global mitochondrial DNA
phylogeography of holarctic breeding dunlins (Calidris alpina ).
– Evol. 50: 318–330.
Wenink, P. W., Baker, A. J. and Tilanus, M. G. 1993.
Hypervariable-control-region sequences reveal global population
structuring in a long-distance migrant shorebird, the Dunlin
(Calidris alpina ). – P. Natl. A. Sci. 90: 94–98.
Wennerberg, L. 2001. Breeding origin and migration pattern of dunlin
(Calidris alpina ) revealed by mitochondrial DNA analysis. – Mol.
Ecol. 10: 1111–1120.
Wennerberg, L., Holmgren, N. M. A., Jönsson, P. E. and Schantz, T. V.
1999. Genetic and morphological variation in Dunlin Calidris
alpina breeding in the Palearctic tundra. – Ibis 141: 391–398.
Weston, M. A., Clarke, K., Maguire, G. S., and Sumner, J. 2020.
Morphological and molecular evidence of population divergence in a
widespread shorebird across its southern mainland Australian
distribution. – Conserv. Genet. 21: 757–770.
Yang, H., Ma, M., Thompson, J. R., and Flower, R. J. 2017. Protect
coastal wetlands in China to save endangered migratory birds. – P.
NatL. A. Sci. 114: 5491–5492.