Acknowledgements
The field work portion of this study was funded by The Negaunee Foundation. We gratefully acknowledge support from Terra Peninsular for facilitating field collections in Reserva Natural Punta Mazo and Enrique Alfaro for his expertise and guidance. We also acknowledge support for the College of Life Sciences and Department of Biology at Brigham Young University and the Grainger Bioinformatics Center at the Field Museum. We thank Jacob Jensen, Zachary Muscavitch, and Emmanuel Serusiaux for fruitful discussions.