Acknowledgement
We are grateful to the Genotoul bioinformatics platform Toulouse Midi-Pyrenees (Bioinfo Genotoul; http://bioinfo.genotoul.fr/) for providing computing resources. We are indebted to Oscar Lao for fruitful discussions and careful reading of the manuscript. This work was supported by two ATM grants (2016 and 2017) from the Muséum National d’Histoire Naturelle to S.M.