Acknowledgements
We are grateful to all the team members in 2012-Transect-Investigation
in the northern grassland of China. We thank Otso Ovaskainen, Nerea
Abrego, Øystein Opedal, and others for organizing the workshop:Multivariate modelling in ecology and joint species
distribution models . This work was supported by National Key Research
and Development Program of China (2016YFC0500703) and Natural Science
Foundation of China (31822006).