Author contributions
K.K.P. and Y.T. did yeast two-hybrid, bimolecular fluorescence complementation, Arabidopsis transformation. K.K.P., Y.T., X.Z.S., Z.P.R. and J.P.X. made the gene cloning, 5’ RLM RACE and DNA sequencing. K.K.P., Y.T. and C.H.S. did western blots and co-express tae-miR399 and MREs in N. benthamiana . K.K.P., Y.T., Y.Z.B., X.Z.S., Y.S.L., J.Y., D.Z. and Q.H.X. measured the plant physiological and phenotypic indexes and analyzed the data. J.C. designed the experiments. K.K.P. and Y.T. wrote the manuscript. All authors discussed the results and approved the final manuscript.