Author contributions
P. Y. and W. Z. conceived and designed the investigation. W. Z., L. N., and L. J. performed field and laboratory work. Z. W. assembled the geome. H. L. performed the Hi-C scaffold. Y. R., H. X., W. Z., Z. W., and H. L. analyzed the data. P. L., P. Y., and W. Z. contributed materials and reagents. P. Y. and W. Z. wrote the paper. All the authors read and approved the final manuscript.