ACKNOWLEDGEMENTS
This study was funded by the National Natural Science Foundation of China (grant number: 31872251, 31500452, and 31372227) and the Earmarked Fund for the Anhui Fishery Research System (2016-84). We acknowledge the support from the Provincial Key Laboratory of Biotic Environment and Ecological Safety in Anhui, and we also are grateful to all colleagues for their performance of field surveys and fish identifications. This research was approved by the Anhui Normal University Animal Ethic Committee. We would like to express our thanks to Gareth Jenkins, Simon Linke, and two anonymous reviewers for their comments, which significantly improved the manuscript.