Acknowledgments

This work was supported equally by the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB31000000), the Second Tibetan Plateau Scientific Expedition and Research (STEP) program (2019QZKK0502), the National Natural Science Foundation of China (grant numbers 31590821 and 91731301), National Key Research and Development Program of China (2017YFC0505203), and, the Fundamental Research Funds for the Central Universities (Grant No. lzujbky- 2019), and International Collaboration 111 Programme (BP0719040). We received support for computational work from the Big Data Computing Platform for Western Ecological Environment and Regional Development and Supercomputing Center of Lanzhou University.