Author Contributions

SUP, SD and SJ designed the study. RDTS, DNS, QC and FH identified study areas and designed fieldwork. SUP, RDTS, DNS, XD, FH, and PBF performed the fieldwork. XD conducted the laboratory work. XD, SD performed the data analysis. XD and AF drafted the manuscript. All authors contributed to the writing of the manuscript. All authors approved the final draft of the manuscript.