References
Data Accessibility and Benefit-Sharing Section: All data used in this manuscript will be made available in the dryad repository upon acceptance for publication.
Author contributions: KS conceptualised, designed and led the research, collected and analysed data, wrote the manuscript; KR contributed to conceptual development and manuscript writing; BKK contributed to generation of phenotype data; JP contributed to designing phenotypic analyses; AE collected field data; RGS generated the phenotype data; MB contributed ecological data; ZOJ contributed to conceptualisation, all molecular work and manuscript writing. All authors contributed to editing the manuscript and approved the final version.