4 Center for Ecological Research, Northeast
Forestry University, 150040 Harbin China
Changjin Cheng
(chengcj@bjfu.edu.cn);
Mingxu Li
(mingxuli@igsnrr.ac.cn);
Congcong Liu
(liucc@igsnrr.ac.cn); Li
Xu (xuli@igsnrr.ac.cn);
*Corresponding authors: Nianpeng He (Institute of Geographic Sciences
and Natural Resources Research, Chinese Academy of Sciences, Beijing,
China; Tel.: +86 10 64889263; Fax: +86 10 64889432; E-mail:
henp@igsnrr.ac.cn)
This manuscript had 150 words in the abstract, 4972 words in the main
text, 55 references, 2 table, and 4 figures.Author contribution
N.H. planned and designed the research; C.C., M.L., C.L. and L.X.
conducted fieldwork and collected data; C.C. analyzed data and wrote the
manuscript; and all authors contributed substantially to revisions.