Acknowledgements
We are thankful to the herbaria of Kew (K) and Seville (SEV) for providing access to their collections for this study and to the colleagues who collected samples in the field. This work was supported by a Marie Skłodowska-Curie Individual Fellowship (704464 – YAMNOMICS – MSCA-IF-EF-ST) and the RBG Kew-funded pilot study to Juan Viruel; and Erasmus+ Programme for Miguel Campos. Emma Kelley received funding from the Muriel Kohn Pokross ’34 Travel/Internship Fund, Smith College, Massachusetts. Pilar Catalán was supported by a European Social Fund and Spanish Aragón Government Bioflora grant.