Affiliations:
1Institute for Environmental Studies, Vrije Universiteit Amsterdam; 1081 HV, Amsterdam, the Netherlands. f.riva@vu.nl2Kellogg Biological Station, Michigan State University; MI 49060, Hickory Corners, Michigan, United States. nick.m.haddad@gmail.com3Geomatic and Landscape Ecology research Laboratory, Carleton University; K1S 5B6, Ottawa, Ontario, Canada. lenorefahrig@cunet.carleton.ca4Department of Life Sciences, Imperial College London; SL5 7PY, Ascot, United Kingdom. c.banks@imperial.ac.uk *Corresponding author. †All authors contributed equally to this work