Acknowledgements
We thank the large number of individuals who have helped with field sampling in the long-term experiment. This research was financially supported by the National Key Research and Development Program of China (2022YFF1302300), the National Natural Science Foundation of China (32260301), the Inner Mongolia Science Fund for Distinguished Young Scholars (2023JQ09, BR230301) and the Local science and technology development fund guided by the central government (2022ZY0210). CMZ was supported by the SNF Ambizione Fellowship program (#PZ00P3_193646).