Parentage analysis
After COLONY’s parentage reassignment, genotype reconstruction, and filtering with the different thresholds, 105 nests were assigned to 27 genotyped mothers, and 13 were assigned to 4 reconstructed females between seasons 2014/15 and 2020/21. The total of 118 nests accounted for 448 offspring. One hatchling that was not assigned to any nest was linked with a genotyped mother. Out of these 118 nests, 70 counted more than one genotyped hatchling. Once the father’s genotypes were reconstructed, a total of 27 fathers explained the genotypes of the 448 hatchlings. The operational sex ratio (OSR) ranged from 0.8 to 1.3 males per female depending on the season, averaging at 0.87, but the sample size was small with fewer than 10 males and females per season. (Table 2). Thus, temporal variations between the nesting seasons were not investigated.