Acknowledgements
This work was funded by the United States National Science Foundation (Award Numbers OCE-1061475, OCE-1450214 and OCE-1623867) to J. D. Witman. We thank L. Dee for assistance in the field, J. Palardy for assistance with preliminary data exploration, boat captains (L. Cruz and W. Aguirre), others (N. Tirado, H. Jager, M. Romoleroux) at the Charles Darwin Foundation and J. Suarez at Galápagos National Park for their assistance. This research was performed under permits from the Galápagos National Park and represents contribution number _______ from the Charles Darwin Foundation for the Galápagos Islands.