Author Contributions
Y.X., W.G. and M.W. contributed equally to this work. Y.X., W.G., X.W., H.W., C.Z., Y.C., M.L. Z.Y., and Z.S. designed the experiments. Y.X., W.G, M.W., X.J., D.H., P.H., M.Z., Q.Z., and P.W. performed the experiments. Y.X., W.G, M.W., X.J., D.H., Z.Y., M.L., G.W. and Y.C. performed the data analysis. Y.X., W.G, M.W., Z.Y., M.L. and Y.C. wrote the manuscript.