Carbon dioxide had significant interactive effects with temperature or light on several oxygen flux parameters. There was a significant interactive effect among temperature, light and CO2 for net photosynthesis per cm2 of coral skeleton and per Symbiodinium cell. However, there was no effect of CO2 alone on the rate of photosynthesis. Godinot (2011) found that ocean acidification of 2242 µatm pCO2 depressed the rate of photosynthesis per µg Chlorophyll a in Stylophora pistillata at 33°C but not 26°C, whereas Crawley et al. (2010) found that photosynthesis per mmol Chlorophyll a (but not per symbiont cell) declined with increasing carbon dioxide. Hii et al. (2009) found that increased carbon dioxide caused decreased (net and gross) photosynthesis and respiration per unit skeletal area in Porites cylindrica and increased respiration and gross photosynthesis (but not net photosynthesis) per unit skeletal area in Galaxea fascicularis. Houlbrèque (2012) found no effect of increased pCO2 on net photosynthesis and respiration normalised to Chlorophyll a in Stylophora pistillata. In a meta-analysis, Harvey et al. (2013) identified a trend for ocean acidification to negatively affect photosynthesis in corals (but the opposite occurred in phytoplankton and seagrass).