Carbon dioxide did not cause a significant change in Chlorophyll a or Chlorophyll c2 per cell, which contrasts some previous studies. Crawley et al. (2010) that found that increased carbon dioxide caused enhanced Chlorophyll a per cell (and thus enhanced photoacclimation) at low light levels. Reynaud et al. (2003) found that increased carbon dioxide combined with higher temperature resulted in increased Chlorophyll a per cell in S. pistillata. Towle et al. (2015) found no effect of pCO2 on Chlorophyll a in Acropora cervicornis.