Temperature had a significant effect on oxygen flux during dark respiration, photosynthesis, and light enhanced dark respiration all standardised to symbiont cell (Figure 2b), surface area, µg of Chlorophyll a and µg of Chlorophyll c2. Temperature increased respiration rates and reduced net photosynthesis at each of these standardisations. Neither light nor pCO2 alone had any effect on oxygen flux per cell or per cm2, and there was no two-way interactive effect between temperature and light, light and pCO2 or temperature and pCO2 (Table 2), except for an interactive effect of pCO2 and temperature on dark respiration per cm2 (p=0.0496) (Figure 2 c).